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Gubler is a high school graduate of the Las Vegas Academy of International Studies, Visual and Performing Arts, where he majored in acting, because the school did not offer his first choice, Filmmaking. He voiced the character of Simon (the smart Chipmunk) in Alvin and the Chipmunks (2007), Alvin and the Chipmunks: The Squeakquel (2009), Alvin and the Chipmunks: Chipwrecked (2011), and Alvin and the Chipmunks: The Road Chip (2015) alongside Jason Lee (Dave), Justin Long (Alvin), and Jesse Mc Cartney (Theodore). He has also provided on/off voice-work for Bagdasarian Productions (regarding the character Simon), whenever Ross Bagdasarian Jr. This is most notable in the episode "Who's The Animal? In 2014, Gubler acted in the zombie comedy Life After Beth and the film Suburban Gothic, for which he received the Screamfest Award for Best Actor.He is a graduate of New York University's Tisch School of the Arts, where he majored in film directing. He appeared as Bart, the main character, in How to Be a Serial Killer (2008), and as Paul in (500) Days of Summer (2009). Gubler has done voice work for the DC Universe Animated Original Movies line.While at NYU film school, Gubler was found by a model scout and then worked as a model with DNA Model Management for Tommy Hilfiger, Marc Jacobs, and American Eagle, After he began modeling, Gubler had an internship with Wes Anderson, who encouraged him to audition for a part in his movie The Life Aquatic with Steve Zissou. He played Jimmy Olsen in All-Star Superman and voices The Riddler in the animated film: Batman: Assault on Arkham (2014).While making The Life Aquatic with Steve Zissou, Gubler made a documentary titled Matthew Gray Gubler's Life Aquatic Intern Journal, a behind-the-scenes view of the making of the film, which was later included in The Criterion Collection DVD release.
Gubler directed the Criminal Minds season 5 episode "Mosley Lane" (5x16), which originally aired March 3, 2010, along with the episode "Lauren" (6x18), which aired a year later on March 16, 2011.The episode "Lauren" included the exit of Criminal Minds series regular Paget Brewster, who maintains a friendship with Gubler outside of work on the show.